Wycliff Palu has overcome an injury cloud to take his place for the Wallabies when they take on Wales on Saturday.
Digby Ioane, Will Genia, Wycliff Palu and Benn Robinson have all been cleared, and today named in the Qantas Wallabies side that will play Wales at the Millennium Stadium on Saturday.
The quartet were all previously in doubt due to injury, with Ioane being a late withdrawal from last weekend's Test against Scotland at Edinburgh, while Genia, Palu and Robinson were forced from the field in that match.
Their roles were confirmed after they each successfully came through training at the University of Glamorgan today.
Ioane's return at centre in place of Ryan Cross is one of three changes to the Australian starting XV from that which took the field in the Hopetoun Cup Test match. The others see flanker David Pocock and lock Dean Mumm gain starts, for George Smith and Mark Chisholm respectively.
Pocock was rested last weekend against Scotland, after being heavily involved earlier in the tour, while Mumm has been rewarded for his all-round efforts.
As well as being a member of the Test match squad through the trip, the NSW Waratahs second rower also led the midweek Australian XV to victories over firstly Gloucester, and then Cardiff earlier this week.
Both Smith and Chisholm have been included on the bench, as has NSW Waratahs utility back Kurtley Beale and Western Force prop Matt Dunning.
Beale, who will make his debut if called onto the field, scored two tries from the unfamiliar wing position during Tuesday night's 31-3 win over Cardiff. He offers additional cover at flyhalf, inside centre, wing and fullback.
Cross, loose forward Richard Brown and prop Sekope Kepu have all been omitted after featuring in last week's match night squad.
Qantas Wallabies coach Robbie Deans says the recovery of the injured quartet is a big boost for his side as it bids to finish a demanding year strongly.
"They're all, in their own right, influential players, both in terms of their actions and the impact that they have on their mates around them, so it is good to have them all back on deck," Deans says.
"No one was more disappointed than us, in terms of the way our most recent Test outing turned out. It was an incredibly frustrating night for everyone involved, as it no doubt was for everyone who was watching back home. We can't alter that outcome now but we're not dwelling on it either. We will learn the lessons from that experience and move on. As a group, we are fortunate that we have another opportunity, so that we can attempt to end this tour, and the year, on a more positive note."
Scotland's 9-8 win last weekend has denied the Wallabies an opportunity to complete the European leg of their expedition unbeaten.
In Wales, they now face an opponent that edged out last year's tourists 21-18, to secure the James Bevan Trophy for the first time.
"This match is a big test of our collective character as a group, there's no doubt about that," Deans says.
"As they showed when we visited last year, and in the recent home matches that they've had against South Africa and the All Blacks, the Welsh are a formidable opponent. They've consistently been one of the best teams in Europe during the last two years, have an experienced playing roster, and had a wide representation on the British Lions tour earlier in the year."
Wales approaches the final game of its November series confident after piecing together successive victories over Samoa and Argentina following a first-up 12-19 defeat by New Zealand.
Just six of the players named in Qantas Wallabies starting line-up announced today were also in the run on side 12 months ago, when Wales beat Australia.
Those players are Ioane, Drew Mitchell, Peter Hynes, Matt Giteau, Stephen Moore and Benn Robinson. Mitchell (who played fullback) and Ioane (wing) are in different positions a year on.
Saturday represents the Wallabies' 14th and final Test of the year, with the majority of the team returning to Australia on Sunday night.
The Qantas Wallabies, is:
15. Adam Ashley-Cooper (Brumbies)
14. Peter Hynes (Queensland Reds)
13. Digby Ioane (Queensland Reds)
12. Quade Cooper (Queensland Reds)
11. Drew Mitchell (NSW Waratahs)
10. Matt Giteau (Brumbies)
9. Will Genia (Queensland Reds)
8. Wycliff Palu (NSW Waratahs)
7. David Pocock (Western Force)
6. Rocky Elsom (Brumbies, captain)
5. Dean Mumm (NSW Waratahs)
4. James Horwill (Queensland Reds)
3. Ben Alexander (Brumbies)
2. Stephen Moore (Brumbies)
1. Benn Robinson (NSW Waratahs)
16. Tatafu Polota-Nau (NSW Waratahs)
17. Matt Dunning (Western Force)
18. Mark Chisholm (Brumbies)
19. George Smith (Brumbies)
20. Luke Burgess (NSW Waratahs)
21. James O'Connor (Western Force)
22. Kurtley Beale (NSW Waratahs)
The Half a Game initiative is currently sweeping across Wales and proving a major success. The scheme aims to give every young player at least half a game every weekend. WRU TV recently visited a festival in Narberth where clubs have fully embraced the initiative.
Both Bargoed and Swansea were seeking their first win of the Principality Premiership season this afternoon. It proved a fruitful day for the hosts - not only did they win, but Bargoed claimed a try bonus point.
Cardiff managed to keep their unbeaten record at home intact, but only just as Bridgend made them work extremely hard before the Blue and Blacks secured a hard fought Principality Premiership win at the BT Sport Arms Park last night.
Former Wales back row Andy Powell gives a guided tour of The Wern, home of Merthyr RFC, who are determined to prove a lot of people wrong and make their mark in their first season in the Principality Premiership.
Wales Sevens head coach Gareth Williams looks back on the Rio Sevens success where he helped GB win silver. He believes the successful campaign is a huge boost to his Welsh squad as they prepare for another World Series, as WRU TV's Graeme Gillespie discovers
British and Irish Lions head coach Warren Gatland will be heading back home to take on the All Blacks next year where for him, success for 2017 is all about going to New Zealand and winning a Test Series.
Last week all roads led to Eugene Cross Park where defending champions Ebbw Vale hosted this season's launch of the Principality Premiership. WRU TV's Graeme Gillespie was there to capture the thoughts of Damien Hudd, captain of Ebbw Vale, and WRU Head of Performance, Geraint John.
Llandovery made it three in a row when they defeated Aberavon in the final of the National Foster's 7s at BT Sport Cardiff Arms Park on Sunday. On another eventful day of sevens, Newport claimed the Plate and the Dragons won the women's crown
Last week saw Cardiff welcome players from around the world for the 21st Golden Oldies Festival. The tournament features players over 35 years of age, and this year's event saw a select few represent the WRU in a 'friendly' against the Wooden Spoon.
Wales Under 18 captain Callum Bradbury is delighted to have seen his side claim a narrow 18-17 victory against a strong SA Schools 'A' side to end the three-match Under 19 International Series in South Africa on a high note.