Wycliff Palu has overcome an injury cloud to take his place for the Wallabies when they take on Wales on Saturday.
Digby Ioane, Will Genia, Wycliff Palu and Benn Robinson have all been cleared, and today named in the Qantas Wallabies side that will play Wales at the Millennium Stadium on Saturday.
The quartet were all previously in doubt due to injury, with Ioane being a late withdrawal from last weekend's Test against Scotland at Edinburgh, while Genia, Palu and Robinson were forced from the field in that match.
Their roles were confirmed after they each successfully came through training at the University of Glamorgan today.
Ioane's return at centre in place of Ryan Cross is one of three changes to the Australian starting XV from that which took the field in the Hopetoun Cup Test match. The others see flanker David Pocock and lock Dean Mumm gain starts, for George Smith and Mark Chisholm respectively.
Pocock was rested last weekend against Scotland, after being heavily involved earlier in the tour, while Mumm has been rewarded for his all-round efforts.
As well as being a member of the Test match squad through the trip, the NSW Waratahs second rower also led the midweek Australian XV to victories over firstly Gloucester, and then Cardiff earlier this week.
Both Smith and Chisholm have been included on the bench, as has NSW Waratahs utility back Kurtley Beale and Western Force prop Matt Dunning.
Beale, who will make his debut if called onto the field, scored two tries from the unfamiliar wing position during Tuesday night's 31-3 win over Cardiff. He offers additional cover at flyhalf, inside centre, wing and fullback.
Cross, loose forward Richard Brown and prop Sekope Kepu have all been omitted after featuring in last week's match night squad.
Qantas Wallabies coach Robbie Deans says the recovery of the injured quartet is a big boost for his side as it bids to finish a demanding year strongly.
"They're all, in their own right, influential players, both in terms of their actions and the impact that they have on their mates around them, so it is good to have them all back on deck," Deans says.
"No one was more disappointed than us, in terms of the way our most recent Test outing turned out. It was an incredibly frustrating night for everyone involved, as it no doubt was for everyone who was watching back home. We can't alter that outcome now but we're not dwelling on it either. We will learn the lessons from that experience and move on. As a group, we are fortunate that we have another opportunity, so that we can attempt to end this tour, and the year, on a more positive note."
Scotland's 9-8 win last weekend has denied the Wallabies an opportunity to complete the European leg of their expedition unbeaten.
In Wales, they now face an opponent that edged out last year's tourists 21-18, to secure the James Bevan Trophy for the first time.
"This match is a big test of our collective character as a group, there's no doubt about that," Deans says.
"As they showed when we visited last year, and in the recent home matches that they've had against South Africa and the All Blacks, the Welsh are a formidable opponent. They've consistently been one of the best teams in Europe during the last two years, have an experienced playing roster, and had a wide representation on the British Lions tour earlier in the year."
Wales approaches the final game of its November series confident after piecing together successive victories over Samoa and Argentina following a first-up 12-19 defeat by New Zealand.
Just six of the players named in Qantas Wallabies starting line-up announced today were also in the run on side 12 months ago, when Wales beat Australia.
Those players are Ioane, Drew Mitchell, Peter Hynes, Matt Giteau, Stephen Moore and Benn Robinson. Mitchell (who played fullback) and Ioane (wing) are in different positions a year on.
Saturday represents the Wallabies' 14th and final Test of the year, with the majority of the team returning to Australia on Sunday night.
The Qantas Wallabies, is:
15. Adam Ashley-Cooper (Brumbies)
14. Peter Hynes (Queensland Reds)
13. Digby Ioane (Queensland Reds)
12. Quade Cooper (Queensland Reds)
11. Drew Mitchell (NSW Waratahs)
10. Matt Giteau (Brumbies)
9. Will Genia (Queensland Reds)
8. Wycliff Palu (NSW Waratahs)
7. David Pocock (Western Force)
6. Rocky Elsom (Brumbies, captain)
5. Dean Mumm (NSW Waratahs)
4. James Horwill (Queensland Reds)
3. Ben Alexander (Brumbies)
2. Stephen Moore (Brumbies)
1. Benn Robinson (NSW Waratahs)
16. Tatafu Polota-Nau (NSW Waratahs)
17. Matt Dunning (Western Force)
18. Mark Chisholm (Brumbies)
19. George Smith (Brumbies)
20. Luke Burgess (NSW Waratahs)
21. James O'Connor (Western Force)
22. Kurtley Beale (NSW Waratahs)
Wales head coach Rob Howley explains his decision to change the captaincy of the national side with Ospreys second row Alun Wyn Jones taking over from Sam Warburton for the forthcoming RBS 6 Nations. Howley is hoping his decision will allow Warburton to rediscover his 'mojo'.
How do you follow a dancing hippopotamus - the Ospreys wheelchair rugby team has just the 'oneness' to do it say the BBC! For a decade the sight of a herd of hippos synchronised swimming around their own private blue lagoon, has lit up our television screens at regular intervals between favourite BBC One shows. But now the nation's most watched channel has relinquished its famous amphibious mammals in favour of a new show-reel which includes the Ospreys wheelchair rugby team.
Head coach Rowland Phillips and key front rower Carys Phillips look ahead to games against Spain and Ireland as Wales Women ramp up their preparations for the forthcoming Six Nations. The two warm-up games are a 'massive opportunity to refine the squad' according to the Welsh camp.
Martyn Phillips has discussed details of the newly published 'Strategy for Welsh Rugby' which puts in black and white terms the plans and direction of the governing body of the national sport in Wales for the next decade.
WRU chief executive Martyn Phillips has urged Welsh rugby fans to follow in the footsteps of their soccer counterparts at last summer's Euro Championship by backing their team to the hilt in the upcoming RBS 6 Nations campaign.
Chief executive Martyn Phillips has highlighted the plans for Wales to become a breeding ground for national and elite club rugby coaches, which are contained in the WRU's 10-year 'Strategy for Welsh Rugby'.
Andrew Coombs is enjoying his new role as Nelson forwards coach. The former Dragons and Wales lock has big ambitions to become a professional coach but feels it's important to learn the ropes in grassroots rugby and where better than his home club Nelson, with friends and family close to hand.
Students at Brynllywarch Hall School, near Newtown, have transformed their own lives, and are now having a positive impact on others through rugby. The sport has had a life-changing effect on the well-being of the students who are now delivering rugby sessions to local primary schools and playing for a new mixed ability side
Wales reached their first semi-final since 2013 in the opening round of this season's HSBC World Rugby Sevens Series in Dubai. WRU TV's Graeme Gillespie recaps on what proved to be an eventful tournament
Wales and Ospreys back row James King, who came through the North Wales age grade system, nominated North Wales Regional Community Rugby Manager Marc Roberts, in the WRU's 'Diolch' initiative where squad players thanked those who played a part in their rise to the top.
Beating Merthyr has boosted Carmarthen Quins hopes of reaching a top eight spot in the Principality Premiership. Quins captain Haydn Pugh is delighted with his team's progress and is determined that everyone from the Under 6s through to the senior team, enjoys the ride.
When the Wales squad were asked to nominate individuals who played a big part in their early playing days, Wales and Dragons full back and wing Hallam Amos thought of his former Monmouth School rugby coach John Bevan.
Sam Cross and Luke Treharne have been named as co-captains of the Wales Sevens side as head coach Gareth Williams names his 12-man squad to play in the first leg of the HSBC World Series in Dubai next weekend.
Bedwas head coach Steve Law and players Nicky Griffiths and Alun Rees extol the virtues of the close knit club as they prepare to host third place RGC 1404 - who are one place head of them on the Principality Premiership points table - on Saturday.