Neath skipper Gareth Llewellyn, the grand daddy of Welsh forwards, is in line to extend his Welsh cap record for a pack member at Twickenham on Saturday.
The 33-year-old has been elevated from the Wales A team to take Ian Gough's place among the replacements following the Newport lock's failure to pass a fitness test.
If Llewellyn makes it onto the field then he will win his 65th cap. He made his debut as a 20-year-old against New Zealand in 1989.
If Wales coach Steve Hansen gives him a run then he will become the longest surving international player in the world today - even beating his former NEC Harlequins team mate, the England prop Jason Leonard, who was first capped in 1990.
A veteran of nine previous games against England, the man who led Wales on seven occasions was a winner against the auld enemy in 1993.
Llewellyn won his last cap in the 1999 World Cup, against Samoa and could aim for a third World Cup in Australia next year.
WRU International Skills Coach Jason Strange is a busy man, combining the new role with that of Wales Under 20 attack coach, Ebbw Vale head coach and continuing to teach at Ebbw Fawr Learning Community.
Warren Gatland and Sam Warburton look ahead to this week's crunch RBS 6 Nations tie against France, where the Wales skipper in particular is keen to make his mark, as Paris has proved a bleak place for him in the past.
Following their historic win over England, Wales Under 20 suffered a narrow, 36-34 loss to their Scottish counterparts last weekend, despite outscoring them by five tries to three. Watch the match highlights and reaction.
WRU TV go behind the scenes with the Wales squad as they put the final touches in place ahead of the clash against Scotland at Murrayfield. Follow the squad on their captain's run, the kickers visit to Murrayfield and Neil Jenkins' press conference