Neath skipper Gareth Llewellyn, the grand daddy of Welsh forwards, is in line to extend his Welsh cap record for a pack member at Twickenham on Saturday.
The 33-year-old has been elevated from the Wales A team to take Ian Gough's place among the replacements following the Newport lock's failure to pass a fitness test.
If Llewellyn makes it onto the field then he will win his 65th cap. He made his debut as a 20-year-old against New Zealand in 1989.
If Wales coach Steve Hansen gives him a run then he will become the longest surving international player in the world today - even beating his former NEC Harlequins team mate, the England prop Jason Leonard, who was first capped in 1990.
A veteran of nine previous games against England, the man who led Wales on seven occasions was a winner against the auld enemy in 1993.
Llewellyn won his last cap in the 1999 World Cup, against Samoa and could aim for a third World Cup in Australia next year.
As the countdown continues for RWC2015, Tournament Chief Executive Debbie Jevans visited the Millennium Stadium to discuss preparations for the eight matches and reveal the positive news that 100,000 tickets for games at the home of Welsh rugby have sold in the last seven days.
Jason Harries is targeting a first win on the World Series circuit this season with Wales. The series returns to action this weekend in Dubai where Harries is part of a team hoping to build from its efforts in round one in October.
Wales skipper Jevon Groves says his side have addressed a few points from Gold Coast and aim to 'hit the ground running' in the next two rounds of the World Series, kicking off with this weekend in Dubai before heading off to South Africa next week.
Wales Sevens manager Steff Thomas is hopeful his side can build from a quarter-final spot in the opening round of the World Series in Australia when his side take part in the forthcoming double header in Dubai and South Africa.