Neath skipper Gareth Llewellyn, the grand daddy of Welsh forwards, is in line to extend his Welsh cap record for a pack member at Twickenham on Saturday.
The 33-year-old has been elevated from the Wales A team to take Ian Gough's place among the replacements following the Newport lock's failure to pass a fitness test.
If Llewellyn makes it onto the field then he will win his 65th cap. He made his debut as a 20-year-old against New Zealand in 1989.
If Wales coach Steve Hansen gives him a run then he will become the longest surving international player in the world today - even beating his former NEC Harlequins team mate, the England prop Jason Leonard, who was first capped in 1990.
A veteran of nine previous games against England, the man who led Wales on seven occasions was a winner against the auld enemy in 1993.
Llewellyn won his last cap in the 1999 World Cup, against Samoa and could aim for a third World Cup in Australia next year.
The last five years have been an emotional rollercoaster on and off the pitch for RGC1404. Promotion to the Principality Premiership was the culmination of five years of hard work and investment in North Wales. WRU TV caught up with prop Joe Simpson, who was one of the first intake of WRU North Wales Academy players in 2010.
Ebbw Vale captain Damien Hudd picked up the biggest accolade of the night at the 11th Principality Premiership Awards Evening held at Principality Stadium, after leading his side to the title for the first time.
Wales assistant coach Robin McBryde is predicting a tough battle against England on Sunday despite the fact they may not be considered to be at full strength. Players will be out to impress Eddie Jones which makes England a dangerous proposition, he says.
Wales U20 captain Tom Phillips and performance psychologist Dr Rich Neil explain why spending three days at Monmouth RFC has further strengthened the togetherness of the squad ahead of the #WorldRugbyU20s