Captain Scott Quinnell will become only the 12th player to win 50 caps for Wales when he leads Wales against England in the Lloyds TSB Six Nations Championship match at Twickenham on Saturday (kick-off 2.30 pm)
The Llanelli captain and No 8, who will be skippering his country for the seventh time, will join Neil Jenkins, Ieuan Evans, Gareth Llewellyn, Garin Jenkins, Gareth Thomas, Rob Howley, JPR Williams, Robert Jones, Gareth Edwards, Scott Gibbs and David Young in the elite '50 club'.
"One of my ambitions was to reach 50 caps for my country," said 29-year-old Quinnell.
"It is an exclusive club with some great players and it is something I want to do."
Wales Coach Steve Hansen has made two changes from the side who beat Italy 44-20 at the Millennium Stadium 18 days ago.
Gareth Thomas is recalled to play in the centre, the Bridgend captain winning his 57th cap as he forms a new midfield partnership with Newport Rugby's Andy Marinos. Thomas comes in for Tom Shanklin.
And in the pack, Llanelli lock Chris Wyatt returns ahead of Ian Gough. Wyatt came on as a replacement against Italy having been injured early on in the opening match of the championship against Ireland in Dublin.
Quinnell, Thomas, Wyatt and scrum half Rob Howley are the only survivors from the starting line-up at Twickenham two years ago.
"After going so close against France, the victory over Italy gave us renewed confidence," said coach Hansen.
"However, England will be smarting from their defeat by France in Paris and we know we face a tough test at Twickenham on Saturday."
With slight injury worries hanging over both Gough and Cardiff utility back Rhys Williams, Hansen has put Gareth Llewellyn and Tom Shanklin on stand-by.
WALES (v England)
Kevin Morgan (Swansea); Dafydd James (Bridgend), Gareth Thomas (Bridgend), Andy Marinos (Newport Rugby), Craig Morgan (Cardiff); Stephen Jones (Llanelli), Rob Howley (Cardiff); Iestyn Thomas (Ebbw Vale), Robin McBryde (Llanelli), Chris Anthony (Newport Rugby), Andy Moore (Swansea), Chris Wyatt (Llanelli), Nathan Budgett (Bridgend), Martyn Williams (Cardiff), Scott Quinnell (Llanelli, captain)
Replacements: Barry Williams (Neath), Spencer John (Cardiff), Ian Gough (Newport Rugby)/Gareth Llewellyn (Neath), Colin Charvis (Swansea), Dwayne Peel (Llanelli), Iestyn Harris (Cardiff), Rhys Williams (Cardiff)/Tom Shanklin (Saracens)
With Liam Williams forced out of the World Cup through injury, lock Luke Charteris and Skills Coach Neil Jenkins say they have to put yesterday's defeat behind them as Wales now look ahead to another physical encounter against South Africa in the quarter-finals of the Rugby World Cup.
WRU Consultant Head of Physical Performance Paul Stridgeon praises the Physical Performance department for their work and dedication as he addresses the media ahead of Wales' Pool A clash against Australia
Twelve young people have been selected to follow a one year WRU Coach Core apprenticeship programme. Coach Core was set up by the Royal Foundation of the Duke and Duchess of Cambridge and Prince Harry as part of the Olympics legacy in 2012 and the Duke of Cambridge was on hand to meet the apprentices on their first day in the job. The programme has been funded by the Hunter Foundation.
The WRU has launched a campaign to create a long term legacy for Welsh club rugby by highlighting the advantages of volunteering. Rhian Edwards, a volunteer at Seven Sisters, has enjoyed many benefits of her volunteering at a grassroots rugby club including being part of the Rugby World Cup volunteer workforce after being nominated by the WRU - and the WRU is asking for more people to develop their 'Welsh rugby roots'.