Captain Scott Quinnell will become only the 12th player to win 50 caps for Wales when he leads Wales against England in the Lloyds TSB Six Nations Championship match at Twickenham on Saturday (kick-off 2.30 pm)
The Llanelli captain and No 8, who will be skippering his country for the seventh time, will join Neil Jenkins, Ieuan Evans, Gareth Llewellyn, Garin Jenkins, Gareth Thomas, Rob Howley, JPR Williams, Robert Jones, Gareth Edwards, Scott Gibbs and David Young in the elite '50 club'.
"One of my ambitions was to reach 50 caps for my country," said 29-year-old Quinnell.
"It is an exclusive club with some great players and it is something I want to do."
Wales Coach Steve Hansen has made two changes from the side who beat Italy 44-20 at the Millennium Stadium 18 days ago.
Gareth Thomas is recalled to play in the centre, the Bridgend captain winning his 57th cap as he forms a new midfield partnership with Newport Rugby's Andy Marinos. Thomas comes in for Tom Shanklin.
And in the pack, Llanelli lock Chris Wyatt returns ahead of Ian Gough. Wyatt came on as a replacement against Italy having been injured early on in the opening match of the championship against Ireland in Dublin.
Quinnell, Thomas, Wyatt and scrum half Rob Howley are the only survivors from the starting line-up at Twickenham two years ago.
"After going so close against France, the victory over Italy gave us renewed confidence," said coach Hansen.
"However, England will be smarting from their defeat by France in Paris and we know we face a tough test at Twickenham on Saturday."
With slight injury worries hanging over both Gough and Cardiff utility back Rhys Williams, Hansen has put Gareth Llewellyn and Tom Shanklin on stand-by.
WALES (v England)
Kevin Morgan (Swansea); Dafydd James (Bridgend), Gareth Thomas (Bridgend), Andy Marinos (Newport Rugby), Craig Morgan (Cardiff); Stephen Jones (Llanelli), Rob Howley (Cardiff); Iestyn Thomas (Ebbw Vale), Robin McBryde (Llanelli), Chris Anthony (Newport Rugby), Andy Moore (Swansea), Chris Wyatt (Llanelli), Nathan Budgett (Bridgend), Martyn Williams (Cardiff), Scott Quinnell (Llanelli, captain)
Replacements: Barry Williams (Neath), Spencer John (Cardiff), Ian Gough (Newport Rugby)/Gareth Llewellyn (Neath), Colin Charvis (Swansea), Dwayne Peel (Llanelli), Iestyn Harris (Cardiff), Rhys Williams (Cardiff)/Tom Shanklin (Saracens)
The Webb Ellis Cup visited Lampeter on Monday to recognise the fact that the town is celebrating a major milestone as it is believed the first ever game of rugby played in Wales, was in fact played in Lampeter 150 years ago.
Cardiff Blues U18 Girls will go into Sunday's Gemau Cymru with an extra spring in their step after Welsh internationals Elinor Snowsill and Philippa Tuttiett offered some last minute tips at a training session earlier this week.
Rob Howley and Josh Lewsey addressed more than 250 community club coaches and referees at the first WRU National Community Conference on the weekend. The volunteers benefitted from coaching masterclasses and workshops in a bid to connect all levels of the rugby pathway in Wales.
UEFA has today confirmed the Football Association of Wales (FAW) has been successful in its bid for the Millennium Stadium to host the 2017 UEFA Champions League Final on Saturday 3rd June, 2017 (kick-off: 19.45hrs GMT).
The winners of a filmmaking competition organised by the education charity Into Film and the WRU enjoyed a behind the scenes tour of the Millennium Stadium - and a chance to be photographed with the WRU's fire-breathing, rugby loving mascot, Scorch The Dragon, and Welsh international, Jake Ball.