Loose-head Jenkins and tight-head Jones suffered long-term injuries but the prop duo are back in action and ready for Twickenham and Jenkins said: "It is great to have Adam back. The last time we played together we started well but we both went off injured.
"It will be nice to get the front-row back together and hopefully have a good forward effort against England."
Wales return to HQ for the first time since breaking their 20-year hoodoo in 2008 after overcoming a 13-point deficit to win 26-19 in Warren Gatland's first match in charge.
That inspired a historic Grand Slam while there has been further success at the venue since with the Blues and Ospreys winning the EDF Energy Cup against English opposition.
Jenkins added: "It makes a difference when you can remember going back into a Twickenham changing rooms knowing that you have celebrated a win.
"To get a win there two years ago was an important achievement for all the boys but it is still a daunting place to go and England are still a very good side at home.
"England have been written off by everyone but we know how good they are and how hard it is to win at Twickenham."